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  • Photophosphorylation – Botany Notes – For W.B.C.S. Examination.

    The third type of phosphorylation to make ATP is found only in cells that carry out photosynthesis.Continue Reading Photophosphorylation – Botany Notes – For W.B.C.S. Examination.

    This process is similar to oxidative phosphorylation in several ways. A primary difference is the ultimate source of the energy for ATP synthesis. In oxidative phosphorylation, the energy comes from electrons produced by oxidation of biological molecules. In photosynthesis, the energy comes from the light of the sun. Photons from the sun interact with chlorophyll molecules in reaction centers in the chloroplasts (Figures <span id="MathJax-Element-1-Frame" class="MathJax" style="font-style: normal;font-weight: normal;line-height: normal;font-size: 17.6px;text-indent: 0px;text-align: left;text-transform: none;letter-spacing: normal;float: none;direction: ltr;max-width: none;max-height: none;min-width: 0px;min-height: 0px;border: 0px;padding: 0px;margin: 0px" role="presentation" data-mathml="5.3.1″>5.3.15.3.1 and <span id="MathJax-Element-2-Frame" class="MathJax" style="font-style: normal;font-weight: normal;line-height: normal;font-size: 17.6px;text-indent: 0px;text-align: left;text-transform: none;letter-spacing: normal;float: none;direction: ltr;max-width: none;max-height: none;min-width: 0px;min-height: 0px;border: 0px;padding: 0px;margin: 0px" role="presentation" data-mathml="5.3.2″>5.3.25.3.2) of plants or membranes of photosynthetic bacteria.

    The similarities of photophosphorylation to oxidative phosphorylation include:

    • a membrane associated electron transport chain
    • creation of a proton gradient
    • harvesting energy of the proton gradient by making ATP with the help of an ATP synthase.

    Some of the differences include :

    • the source of the electrons – H2O for photosynthesis versus NADH/FADH2 for oxidative phosphorylation
    • direction of proton pumping – into the thylakoid space of the chloroplasts versus outside the matrix of the mitochondrion
    • movement of protons during ATP synthesis – out of the thylakoid space in photosynthesis versus into the mitochondrial matrix in oxidative phosphorylation
    • nature of the terminal electron acceptor – NADP+ in photosynthesis versus O2 in oxidative phosphorylation.

    In this process, light energy is captured and used for converting carbon dioxide and water into glucose and oxygen gas. The complete process of photosynthesis is carried out into two processes:

    Light reaction

    The light reaction takes place in the grana of the chloroplast. Here, light energy gets converted to chemical energy as ATP and NADPH. In this very light reaction, the addition of phosphate in the presence of light or the synthesizing of ATP by cells is known as photophosphorylation.

    Dark reaction

    While in the dark reaction, the energy produced previously in the light reaction is utilized to fix carbon dioxide into carbohydrates. The location where this happens is the stroma of the chloroplasts.

    Photophosphorylation

    Photophosphorylation is the process of utilizing light energy from photosynthesis to convert ADP to ATP. It is the process of synthesizing energy-rich ATP molecules by transferring the phosphate group into ADP molecule in the presence of light.

    Photophosphorylation is of two types:

    • Cyclic Photophosphorylation
    • Non-cyclic Photophosphorylation

    The photophosphorylation process which results in the movement of the electrons in a cyclic manner for synthesizing ATP molecules is called cyclic photophosphorylation.

    In this process, plant cells just accomplish the ADP to ATP for immediate energy for the cells. This process usually takes place in the thylakoid membrane and uses Photosystem I and the chlorophyll P700.

    During cyclic photophosphorylation, the electrons are transferred back to P700 instead of moving into the NADP from the electron acceptor. This downward movement of electrons from an acceptor to P700 results in the formation of ATP molecules.

    Non-Cyclic Photophosphorylation

    The photophosphorylation process which results in the movement of the electrons in a non-cyclic manner for synthesizing ATP molecules using the energy from excited electrons provided by photosystem II is called non-cyclic photophosphorylation.

    This process is referred to as non- cyclic photophosphorylation because the lost electrons by P680 of Photosystem II are occupied by P700 of Photosystem I and are not reverted to P680. Here the complete movement of the electrons is in a unidirectional or in a non- cyclic manner.

    During non-cyclic photophosphorylation, the electrons released by P700 are carried by primary acceptor and are finally passed on to NADP. Here, the electrons combine with the protons – H+ which is produced by splitting up of the water molecule and reduces NADP to NADPH2.

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