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  • Mollusca – Zoology Notes For – W.B.C.S. Examination.
    Posted on June 4th, 2019 in Zoology
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    Mollusca – Zoology Notes For – W.B.C.S. Examination.

    কম্বোজ – প্রাণিবিদ্যা নোট – WBCS পরীক্ষা।

    Mollusk, also spelled mollusc, any soft-bodied invertebrate of the phylum Mollusca, usually wholly or partly enclosed in a calcium carbonate shell secreted by a soft mantle covering the body. Along with the insects and vertebrates, it is one of the most diverse groups in the animal kingdom, with nearly 100,000 (possibly as many as 150,000) described species. Each group includes an ecologically and structurally immense variety of forms: the shell-less Caudofoveata; the narrow-footed gliders (Solenogastres); the serially valved chitons (Placophora or Polyplacophora); the cap-shaped neopilinids (Monoplacophora); the limpets, snails, and slugs (Gastropoda); the clams, mussels, scallops, oysters, shipworms, and cockles (Bivalvia); the tubiform to barrel-shaped tusk shells (Scaphopoda); and the nautiluses, cuttlefishes, squids, and octopuses (Cephalopoda).Continue Reading Mollusca – Zoology Notes For – W.B.C.S. Examination.

    Size range and diversity of structure

    Typical molluscan features have been substantially altered, or even lost, in many subgroups. Among the cephalopods the giant squids (Architeuthis), the largest living invertebrates, attain a body length of eight metres (more than 26 feet); with the tentacle arms extended, the total length reaches to 22 metres. Other cephalopods exceed a length of one metre. Many of the remaining molluscan classes show a large variation in size: among bivalves the giant clam (Tridacna) ranges up to 135 centimetres (four feet) and the pen shell (Pinna) from 40 to 80 centimetres; among gastropods the sea hares (Aplysia) grow from 40 to 100 centimetres and the Australian trumpet, or baler (Syrinx), up to 60 centimetres; among placophores the gumshoe, or gumboot chiton (Cryptochiton), achieves a length up to 30 to 43 centimetres; and, among solenogasters, Epimenia reaches a length of 15 to 30 centimetres. Finally, gastropods of the family Entoconchidae, which are parasitic in echinoderm sea cucumbers, may reach a size of almost 1.3 metres. In contrast, there are also minute members, less than one millimetre (0.04 inch) in size, among the solenogasters and gastropods.

    Distribution and abundance

    The mollusks have adapted to all habitats except air. Although basically marine, bivalves and gastropods include freshwater species. Gastropods have also adapted to land, with thousands of species living a fully terrestrial existence. Found on rocky, sandy, and muddy substrata, mollusks burrow, crawl, become cemented to the surface, or are free-swimming.

    Caudofoveates (subclass Chaetodermomorpha, class Aplacophora) burrow in muddy sediments at depths of 10 to more than 7,000 metres (33 to 23,000 feet) and consume microorganisms and loose organic material (detritus). In contrast, solenogasters (subclass Neomeniamorpha, class Aplacophora) prey on some members of the class Cnidaria (e.g., hydroids and corals) in five to 6,850 metres of water, on clay or muddy sand, or directly upon hydroid or coral colonies. Chitons (class Polyplacophora) cling to hard bottoms of the intertidal zone, scraping algae from the rock surfaces by using their strong rasping teeth (radula); several members of the polyplacophoran family Lepidopleuridae consume detritus found at depths down to 7,000 metres, and Hanleyidae as well as Hopaliidae even depend on animal food. The few extant members of the class Monoplacophora inhabit secondary hard bottoms at depths of 175 to 6,500 metres and capture detritus by means of head appendages (velum) around the mouth. The scaphopods dwell in sand or sandy mud down to 7,000 metres and nourish themselves on protozoa, crustaceans, or small mollusks captured by the filamentous head tentacles (captacula). Except for the carnivorous septibranch anomalodermata, all bivalves are ciliary suspension feeders, using food-sorting organs near the mouth (labial palps) and respiratory gills modified to assist in feeding (ctenidia). Found in marine and fresh water, most bivalves burrow into sediments to depths of 10,700 metres or attach themselves to hard surfaces by means of tough threads secreted by the byssus gland in the foot. The members of some species may even bore into wood or rock. Cephalopods are generally carnivores, feeding on crustaceans and fishes, but some have adapted a microvorous diet of detritus and microscopic organisms and plants. Some cephalopods are offshore (pelagic) jet swimmers, moving from the surface to depths of 5,400 metres, while others dwell near the bottom (benthic) at depths of 8,100 metres.The greatest ecological diversity is shown by the gastropods. The marine members are found from the spring-tide line to deep-sea trenches (10,500 metres deep) and inhabit nearly all possible habitats, even floating weeds. Both shelled and naked gastropods have pelagic members that spend their entire lives swimming in the water; others penetrate marine hot vents or interstices between sand grains. Some gastropods are parasitic, while others are predatory. Freshwater snails also are found in groundwaters and may inhabit hot springs. Widely distributed throughout all terrestrial habitats, various members of the gastropod order Stylommatophora are adapted to certain regions.

    Some littoral bivalves, such as Tridacna, as well as some sea slugs, such as Aeolidia, share an obligatory symbiosis with zooxanthellae (a group of algae). Another metabolic association exists between certain bacteria and several bivalves and gastropods of deep-sea hot vents or other sulfidesystems. There are several parasitic mollusks.

    Locomotion

    Mollusks have a wide range of locomotory patterns. Solenogasters and various smaller gastropods glide upon cilia that beat rapidly against a pathway of mucus secretions. This pattern of movement is supported or replaced in larger mollusks by the propulsive waves that run along the surface of the foot and are controlled by the actions of the dorsoventral musculature (Figure 1). Burrowing occurs as an interaction between musculature and the hydrostatic skeleton (see below Internal features); it is performed in caudofoveates and several sea slugs by the whole anterior body but is restricted to the foot in scaphopods, bivalves, and some specialized gastropods.

    Various bivalves (e.g., cockles) and snails may perform rapid twists or jumps through violent flexion of the foot. Buoyancy floating and jet propulsion are found in cephalopods; floating is also known in gastropods, and swimming of a different kind is practiced by some opisthobranch and prosobranch gastropods as well as in scallops and related bivalves. Octopods use their arms to crawl or even to swim or float with the help of the body skin interconnecting the arms (interbrachiate web). Some bivalve groups bore into hard surfaces by secreting strong chemicals that dissolve the substrate or by drilling, using the shell and radula. A sedentary (sessile) way of life has been adopted by many bivalves and some gastropods.

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